Maths is Toothpaste - Mathematics is useless on its own, it only has a function when used with something else (i.e. a toothbrush). An online discussion. - http://mathsistoothpaste.blogspot.com
The Math Forum - Help and discussions about mathematics and philosophy, with problems of the week. - http://www.themathforum.com/
Ars Mathematica - News and discussions on the mathematical arts. - http://www.arsmathematica.net/
Math Help Forum - A discussion forum for students and teachers of mathematics at school, college and advanced levels, including homework help for students. - http://www.mathhelpforum.com/
The CTK Exchange Forums - Discussions on elementary, middle and high school mathematics and mathematics education. - http://www.cut-the-knot.org/cgi-bin/dcforum/ctk.cgi
Math is Fun Forum - A discussion forum for K-12 math students and teachers, with a focus on learning and teaching math the fun way. - http://www.mathsisfun.com/forum/
Math Forum: Public Discussions - Group topics include general math, education, geometry, software, calculus reform, research and recreations. - http://mathforum.org/kb/forumindex.jspa
Mathforge - News and breakthroughs in the mathematical sciences, including software, education and books. - http://mathforge.net/
Undernet Math - Topica email list serving the users of the IRC channel #math on the Undernet, for discussion and challenging puzzles. - http://lists.topica.com/lists/undernet-math/
JISCmail - Mailing lists in mathematics and statistics for the UK academic community. - http://www.jiscmail.ac.uk/mailinglists/category/Maths_&_Statistics.htm
WebMath - A mailing list for those interested in mathematics on the World Wide Web. - http://west.camel.math.ca/cgi-bin/wcms/webmath.pl
Javamath-discuss - A general mailing list for discussion about mathematical software development in Java. - http://lists.sourceforge.net/lists/listinfo/javamath-discuss
Maths-Education - Email forum for those working in mathematics education to share ideas and experiences, and request information. - http://lists.nottingham.ac.uk/mailman/listinfo/maths-education
Yahoo Groups: Theory-edge - Informal discussion of research in algorithmics, mathematics, and their applications. - http://groups.yahoo.com/group/theory-edge/
Yahoo Groups : maththeorists - Discussions relating to number and mathematics philosophy. - http://groups.yahoo.com/group/maththeorists/
Yahoo Groups : Erdos - A group for proposing problems, sharing ideas, discussing areas of research and reviewing papers and books of interest. [Requires Yahoo! membership] - http://groups.yahoo.com/group/Erdos/
Yahoo Groups in Mathematics - Mailing lists and discussion groups (including former OneList and eGroups). - http://dir.groups.yahoo.com/dir/Science/Mathematics
Yahoo Groups: Math For Fun - Focuses on solving challenging math problems. - http://groups.yahoo.com/group/mathforfun/